reproducing, the 'social contract' is destroyed and the colony cohesion is dissolved. In order to minimize these losses, animals generally retreat from fighting or displaying fighting ability unless there are obvious cues indicating victory. (eds) Physiology, vol. Dubai One is expected to reach at least 700m and be completed by metres (2,170 ft) 115 The Burj 2020 will be the largest commercial tower in the world, surpassing the just-built One World Trade Centre in New York. Assuming their high rank is correlated with higher fitness and fighting ability, this trait will be conferred to their offspring. 1, in a despotic system, one member is considered dominant while all other members of the living group are equally submissive.